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Mario, Luigi, Toad, and Yoshi battle Bowser Jr.'s sinister plans to conquer the islands, encountering new foes.
Hint: In a race across the US heartland, a red car discovers the true meaning of friendship.
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The brothers return and face the new arising menace that is Bowser Jr. Mario, Luigi, Toad and Yoshi must stop Bowser and his son from concurring the islands and face new foes along the way